Primus
Primus is Unicron's sibling and the creator of the Transformers in the comic continuity. His body is the planet Cybertron. Primus' lifeforce (aka the Allspark) is a multiversal singularity; it exists across nearly all Transformers realities(Primus is never stated to exist in the cartoon, nor anything resembling Primus).
In the beginning
(The Primus/Unicron backstory has evolved and been rewritten a number of times since it originated in the Marvel Comics. The version below is the current iteration, which has slowly solidified across a wide swath of media begining around the time of Armada)
At the birth of the Universe, an extradimensional entity known as The One created twin astral beings to reign over the cosmos. One was Primus, the Lord of Light and Order. The other was Unicron, the Lord of Darkness and Chaos. Unicron sought to devour everything that exists and become one with The Void at the end of all things.
Primus was no match for Unicron. But Primus did manage to trap both his and Unicron's essences inside metallic planetoids. Over time, they learned to shape their prisons. Primus became the mechanical planet of Cybertron. Knowing that his ancient enemy still lived, Primus created the 13 Original Transformers from the material of the newly created planet. To the very first of these 13, Primus bestowed the Matrix, his essence. These Transformers would, in time, grow strong in number and knowledge and power and be able to defeat Unicron.
The two brothers shared a mental link. So long as Primus hid himself in slumber, so did Unicron also sleep. Their war ended, for a time, in stalemate.
War of the balance
Eventually in one version of Generation 1 continuity Grimlock, Jazz, and Bumblebee found themselves transported to the center of Cybertron. In a battle with Bludgeon's Pretenders, a laser blast ricocheted off Grimlock and struck the essence of the sleeping Primus, causing a multiverse-shaking Primal Scream that woke Unicron and alerted him to Primus' location.
The battle between Primus and Unicron began anew. Though Unicron would sometimes be defeated, this is always temporary, and there is a whole multiverse of realities to consume. When Unicron is destroyed in one reality, he will respawn in another, ready to eat again. Primus will always be waiting, hoping to incapacitate Unicron and preserve existence.
Alternate Backstories
Before the Primus/Unicron Mythos reached its present form it went through several versions in the Marvel G1 Comics where it originated.
The first time we ever heard the Primus/Unicron backstory it was from Unicron himself. After his destruction during the events of Transformers: The Movie his head floated through space and crashed on the planet Junk. There he enslaved the inhabitants to recreate his old body from their world. When the bount-um... freelance peacekeeping agent Death's Head challenged him on the astral plain Unicron shared the story of his origin:
When, at the dawn of the universe, Unicron was a primal force of evil the only one who could stand against him and his Dark Armies was Primus, Lord of the Light Gods. Primus and his fellow Light Gods stood against Unicron and his Dark Armies. Events then played out as described above. (The role of the other Light Gods and Unicron's armies would diminish with each retelling of the story, until the current version where they are alone and have a unique origin. Only one of the other Light Gods was ever named: the Chronarchitect.)
The second time the story was told was by The Keeper an ancient mechanoid who guarded Primus's head at the center of Cybertron. This telling mentions that their battle was towards the end of the era of Gods and that Primus and Unicron were the last of their respective pantheons. But Primus had to defeat Unicron before he could take his place with the other gods in the "Omniversal Matrix".
The third time the story was told was by Primus when he gathered all his children togather to prepare for Unicron's comming. Here we learn for the first time that Unicron predated the current universe. Unicron had destroyed the previous universe and slept peacefully, alone with the void. But he had not been thurough enough and some bits of the "old realms" survived. They reacted and caused The Big Bang. The sentient core of this new universe recognized the threat of Unicron and so created Primus to be his counterpart and guardian of the new creation.
(Primus did not exist in the Generation 1 Cartoon. There, Cybertron's oldest revealed status was as a factory world of the Quintessons who invented the Transformers, and Unicron was a product of the strange sad little alien Primacron.)
Marvel Comics Continuity
Unicron is first seen in the comic adaptation of Transformers: the Movie. The Marvel US comic basically left the adaptation alone as something that might or might not happen but was in the distant future (at the time) and so basicaly out of continuity. The Marvel UK version of the comic however seized on the Movie era and its characters as something it could focus on without worry of contradicting events in the US stories it reprinted.
Target 2006 was the first of these UK exclusive stories. It is not until the later "Legacy of Unicron" storyline however that we first hear of and see Primus.
Beast Wars/Beast Machines
Primus' name was often invoked in prayer or in righteous anger by the Beast Warriors.
The Covenant of Primus, a prophetical set of data tracks, were on board both the Ark and the Nemesis when they crashed 4 million years ago on prehistoric Earth. Megatron himself took his name from these tracks (as did, presumably, the original Megatron), and the scriptures foretold the events which completed the Beast Wars.
After the Beast Wars, Megatron intended to incorporate Primus' essence, the Allspark, into his own being and become godlike by stealing every Spark of every Transformer on Cybertron. Ultimately, this was merely one step of the Oracle's plan to reformat Cybertron into a technorganic paradise. After Megatron and Optimus Primal were plunged into the planet's core, the planet was reborn and the abducted Sparks were returned to new technorganic bodies.
Unicron Trilogy
Energon
Cybertron
Primus' plot to bury Unicron's essence in the center of a newborn star failed. The star collapsed into a black hole which threatened not only Cybertron and the universe, but the multiverse as well. Only four Cyber Planet Keys, hidden on planets throughout the galaxy, combined with the Omega Lock, could awaken Primus and allow him to destroy the black hole.
Universe/Timelines
Back at the dawn of the Universe, before he put into effect his grand plan, Primus did a test run to make sure it would work. He transformed a small moon into a planet and there created twelve transforming, robotic beings based on the Zodiac of future Earth. (He was a forward thinking God and had the Chronarchitect on his side.) The Covenant of Primus as they were called, were given the duty of watching over their younger cousins the Transformers, ensuring that Primus' plan followed its intended path, and preparing for the Omega Point, a prophesized final battle with Unicron in which they were to take part.
When Unicron began gathering Transformers from the multiverse, forcing them to fight, and harvesting the Sparks of the losers, this presented a huge threat to Primus and the Allspark. Transformers' Sparks are pieces of Primus' essence, so each devoured soul weakened him.
Primus and Alpha Trion called back from the Allspark the now-legendary Maximal ship commander and Cybertron's savior, Optimus Primal, to rescue the possessed, battling captives from the Pit of Unicron -- and to lead a team of multiversal warriors to put a stop to his soul-stealing scheme before it could cause the Allspark to collapse.
Primus allowed Optimus Primal to choose any Transformer who ever existed -- living or dead -- for his army. This required considerable amounts of Primus' power, so Primal was told to choose sparingly. Primal chose his departed former troops Rhinox and Depth Charge.
(This storyline folded into the Timelines story from the TF fanclub.)
During the events of Transformers: Cybertron, the black hole created by Unicron's passing posed unique problems. Primus and Unicron -- representing Order and Chaos, respectively -- together forged a delicate balance in the multiverse. With one or the other missing, the balance would be upset, spiralling all that exists into an unknown doom.
In addition, the black hole was positioned very close to Cybertron itself, making Primus, despite Unicron's apparent demise, the most vulnerable he had ever been. Cybertron was the linchpin, the astral center of the entire multiverse universe, itself another singularity. Unicron's heralds Ramjet and Nemesis Prime arrived on Cybertron to unleash the Dead Matrix on Primus' essence. For the first time ever, Unicron, who is normally limited to threatening Primus one reality at a time, would be able to weaken Primus across all timelines -- and the black hole's proximity would end their cosmic feud forever.
Toys
To date, only two toys of Primus have been released throughout the world. The first was a "Lucky Draw" toy in Japan, a recolored metallic Rodimus Prime that was given away as a contest prize from a collector book.
The second was released in Galaxy Force as "Primus" in Spring 2006, only in Japan. The Western version will be released as "Cybertron Primus" in Cybertron from Hasbro, and the first run will include a bonus accessory of Unicron's decaying severed head, a reference to Transformers:The Movie, where Unicron's head orbitted Cybertron.
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