Cosmitron

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So vital of a part that Ironhide can't even recall the name.

A cosmotron is a piece of Cybertronian technology of unspecified function. What is known is that Optimus Prime's cosmotron was severely damaged during a battle with the Decepticons, and the Autobot leader surely would have perished if his cosmotron had not been replaced, suggesting that whatever purpose it serves is a vital one.