Domitius Major

Warp factor four, Mr. Centurion - engage!

Domitius Major is a Eukarian Autobot from the IDW portion of the Generation 1 continuity family.

Domitius Major is a Eukaran on a mission from GodOnyx Prime, commanding an rather familiar ship. After awhile he starts to look a lot like this guy.

He may have punched out Cthulhu.

Fiction

Spoiler warning: Plot details for Strange Visitors follow. Spoilers expire in 36 days

IDW Generation 1 continuity

"And Trix are for protoforms."

Twenty thousand years ago, Dominitus Major and a crew of Eukarian explorers captained a deep space expedition aboard the Axalon at the behest of his leader Onyx Prime. The ship's crew was entrusted with the supernaturally-empowered Talisman. Depleting energon stocks aboard their vessel prompted Domitius to change course for the nearest planet with energon—Earth. As they prepared to set down, however, they first received a transmission of Optimus Prime and Bumblebee—alerting them to Shockwave's own ship, who shot down the colonists.

The Axalon crashed on a glacier in the land that would one day be Brasnya; though physically unharmed, the Talisman their vessel carried had scrambled the memories of their crew, leaving them with no trace of their memories. Centurion's confusion over his lost identity inspired Shockwave—who had originally come to Earth to investigate the progress of his Regenesis project—to use the amnesiac Transformers in a twisted experiment of his own devising. By overwriting their memories with data from the Cybertronian Great War, the amoral scientist was able to convince the stranded explorers that they were Autobots and Decepticons, and Domitius Major was conditioned into believing that he was Optimus Prime. The two factions battled one another for a century in a series of simulated war games while Shockwave observed their tactics. Eventually, however, the Eukarians ran low on energy and entered stasis lock underground. Strange Visitors

In the 19th century, human activity prompted several of the Eukarians to awaken, though "Optimus Prime" was not one of them. Several of the ersatz Autobots mulled over the possibility of reactivating their commander during the brief conflict between the "Autobots" and "Decepticons" in the 19th century, though they ultimately decided against it. Hearts of Steel #1 Hearts of Steel #2

GIVE ME YOUR F...er, tentacles.

Several years later, however, the forces of the Elder Gods manifested themselves, threatening both humans and Transformers. With the help of Tobias Muldoon and Nikola Tesla, the Eukarian Autobots were able to reactivate their commander, who had been buried beneath the Mare Rurek Glacier. "Optimus" battled the Elder God in Nova Scotia, and eventually drove it and its followers back to the depths of the ocean. In the aftermath, he pledged to defend Earth if the abomination ever resurfaced. Infestation 2: The Transformers #2

Some time after that, Domitius and Centurion, still believing themselves to be Optimus and Bumblebee, were caught in a World War I battlefield. Faced with a real battleground, their artificial memories of the Cybertronian war swiftly unraveled, and Domitius perished that day. His death was enough to shake "Bumblebee" from his conditioning.

By the end of World War II, Centurion returned to the crash site of the Axalon, where he discovered that the Iron Troopers that the Talisman created had banded together to puppeteer Domitius's dead body like a suit. Centurion—by this time, the last surviving member of the Axalon crew—battled his old commander for the last time and eventually managed to tear his body apart. Strange Visitors

Domitius's frozen remains were later retrieved by Doctor X, where they were relocated and stored in a secure facility. The Iron Klaw