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In most universal streams, Chromedome is a comparatively quiet academic. Originally a number-cruncher at Cybertron's Institute for Higher Programming working on ways to improve the processing power of the Transformers' computerized minds, Chromedome found himself forcibly dragged into the Great War when a Decepticon attack destroyed the facility. Chromedome put his work to use programming Autobots so they could outsmart their Decepticon foes, but alas, only wound up fostering an intellectual arms race of sorts, in which both sides competed to repurpose the newest advances in programming into military advantages. Ground down over the years by this inevitability, Chromedome has come to accept that war with the Decepticons cannot be prevented—but only won. Well-versed in the mechanisms of the Cybertronian mind, Chromedome has been known to specialize in mnemosurgery, the science of manipulating the Transformer brain and the memories within.

In some universes, Chromedome is binary bonded to the egotistical Nebulan Stylor, who is more concerned with the finer things in life than intellectual or military pursuits. As such, the two occasionally clash, with Chromedome trying to get Stylor to be a little less superficial, and Stylor trying to bring out Chromedome's more social side, but generally speaking, they get along quite well despite their differences.

In other universes, Chromedome's an ex-copper and quiet academic with burning passions. He falls easily in love and even more easily into really bad ideas and self destructive habits. But he can totally quit mnemosurgery any time, guys! He just has a reason why this one time, it's okay.

He's Conjunx Endura with Rewind. Deep down, Chromedome's sure Rewind would leave him for his previous partner.

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